Flora of Newfoundland and Labrador
Flora of Newfoundland and Labrador
Lycopodiaceae/Lycopodiacées - Clubmoss Family
Spinulum annotinum (L.) A.Haines subsp. dubium (Zoëga) J.H.Xue, Chepinoga & K.P.Ma
En: northern bristly clubmoss, northern interrupted clubmoss
Fr: lycopode innovant du Nord, lycopode interrompu du Nord
IU: siqpiijautit, sippiKautik
Provincial Status: L
Range: Circumboreal; Lab., QC-MB, AB-BC, NU-YK; acid.

[=Lycopodium alpestre (Hartm.) Schmakov & Tichonov]
[=Lycopodium annotinum L. forma brevifolium Christ]
[=Lycopodium annotinum L. forma pungens (Desv. ex Spring) Porsild]
[=Lycopodium annotinum L. subsp. alpestre (Hartm.) Á.Löve & D.Löve]
[=Lycopodium annotinum L. subsp. dubium (Zoëga) Kallio, Laine & Y.Mäkinen]
[=Lycopodium annotinum L. subsp. pungens (Desv. ex Spring) Hultén, nom. inval.]
[=Lycopodium annotinum L. var. acicularis Christ]
[=Lycopodium annotinum L. var. alpestre Hartm.  (basionym)]
[=Lycopodium annotinum L. var. pungens Desv ex Spring]
[=Lycopodium annotinum L. var. pungens (Desv.) Bach.Pyl. ex Desv. 1827, nom. inval.]
[=Lycopodium canadense Romhild ex Nessel]
[=Lycopodium dubium Zoëga]
[=Lycopodium neopungens H.S.Kung & Li Bing Zhang]
[=Lycopodium pungens (Desv. ex Spring) Bach.Pyl. ex Iljin 1934, nom. illeg. hom., non Alderw. 1915]
[=Lycopodium subarcticum V.N.Vassil.]
[=Lycopodium zonatum Ching]
[=Spinulum annotinum (L.) A.Haines subsp. alpestre (Hartm.) Uotila]  
[=Spinulum canadense (Romhild ex Nessel) A.Haines]
[=Spinulum subarcticum (V.N.Vassil.) A.Haines] 
[=Spinulum zonatum (Ching) Mazumdar]

[Although not recognized in FNA, this species is usually treated in Europe, as well as the PAF as having two subspecies. The typical subspecies occurs in the more southern range, while subsp. alpestre (Hartm.) Uotila occupies more northern (arctic) areas. A broad hybrid zone occurs where these two subspecies meet and material from this area, including Newfoundland and Labrador, Greenland, and Kamchatka, was described as L. annotinum var. pungens Desv. ex Spring (T. Ahti, pers. comm., April 2015; Elven et al. 2011). Note that the authority of var. pungens should be cited as Desv. ex Spring, since the name in Desvaux's original 1827 publication was an invalid nomen nudum. The name L. annotinum var. pungens Bach.Pyl. ex Desv. (1827) is also invalid, as this name was listed in synonymy; Bachelot de la Pylaie did not make a valid publication in this taxon, so combinations with Bach.Pyl. listed as the basionym are in error.]

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