Flora of Newfoundland and Labrador
Flora of Newfoundland and Labrador
Lycopodiaceae/Lycopodiacées - Clubmoss Family
Spinulum annotinum (L.) A.Haines
En: stiff clubmoss, bristly clubmoss, interrupted clubmoss
Fr: lycopode innovant, lycopode interrompu
IA: kakauat
IU: sippiKautik
Provincial Status: LN
Range: Circumboreal; thr. Nfld. & Lab.; acid.

[=Lepidotis annotina (L.) P.Beauv.]
[=Lycopodium annotinum L.  (basionym)]
[=Lycopodium annotinum L. forma proliferum Lepage]
[=Lycopodium annotinum L. forma pungens (Desv. ex Spring) Porsild]
[=Lycopodium annotinum L. forma substipitatum Luerss.]
[=Lycopodium annotinum L. subsp. alpestre (Hartm.) Á.Löve & D.Löve]
[=Lycopodium annotinum L. subsp. annotinum]
[=Lycopodium annotinum L. subsp. dubium Kallio, Laine & Makinen]
[=Lycopodium annotinum L. subsp. pungens (Bach.Pyl.) Hultén, comb. inval.]
[=Lycopodium annotinum L. subsp. pungens (Desv. ex Spring) Hultén]
[=Lycopodium annotinum L. var. acrifolium Fernald]
[=Lycopodium annotinum L. var. alpestre Hartm.]
[=Lycopodium annotinum L. var. annotinum]
[=Lycopodium annotinum L. var. montanum Tuck.]
[=Lycopodium annotinum L. var. pungens Desv., nom. inval., nom. nud.]
[=Lycopodium annotinum L. var. pungens Bach.Pyl. ex Desv., nom. inval., pro syn.]
[=Lycopodium annotinum L. var. pungens Desv. ex Spring]
[=Lycopodium annotinum L. var. subpungens Macoun ex Nessel, nom. inval., pro syn.]
[=Lycopodium annotinum L. var. tenuifolium Nessel]
[=Lycopodium bryophyllum C.Presl]
[=Lycopodium canadense Nessel]
[=Lycopodium canadense Romhild ex Nessel 1939, nom. inval.]
[=Lycopodium dubium auct., non Zoëga]
[=Lycopodium juniperifolium Lam., nom. illeg.]
[=Lycopodium neopungens H.S.Kung & L.B.Zhang]
[=Spinulum annotinum (L.) A.Haines]
[=Spinulum canadense (Nessel) A.Haines]

[Although not recognized in FNA, this species is usually treated in Europe, as well as the PAF as having two subspecies. The typical subspecies occurs in the more southern range, while subsp. alpestre (Hartm.) Á.Löve & D.Löve occupies more northern (arctic) areas. A broad hybrid zone occurs where these two subspecies meet and material from this area, including Newfoundland and Labrador, Greenland, and Kamchatka, was described as L. annotinum var. pungens Desv. ex Spring (T. Ahti, pers. comm., April 2015; Elven et al. 2011). Note that the authority of var. pungens should be cited as Desv. ex Spring, since the name in Desvaux's original 1827 publication was an invalid nomen nudum. The name L. annotinum var. pungens Bach.Pyl. ex Desv. (1827) is also invalid, as this name was listed in synonymy; Bachelot de la Pylaie did not make a valid publication in this taxon, so combinations with Bach.Pyl. listed as the basionym are in error.]

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